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In 1959, the heart of a 22 metric-ton (24 short-ton) male taken by whalers was measured to be 116 kilograms (256 lb), about 0.5% of its total mass.[79] The circulatory system has a number of specific adaptations for the aquatic environment. The diameter of the aortic arch increases as it leaves the heart. This bulbous expansion acts as a windkessel, ensuring a steady blood flow as the heart rate slows during diving.[80] The arteries that leave the aortic arch are positioned symmetrically. There is no costocervical artery. There is no direct connection between the internal carotid artery and the vessels of the brain.[81] Their circulatory system has adapted to dive at great depths, as much as 2,250 metres (7,382 ft)[6][7][82][83][84][excessive citations] for up to 120 minutes.[85] More typical dives are around 400 metres (1,310 ft) and 35 minutes in duration.[38] Myoglobin, which stores oxygen in muscle tissue, is much more abundant than in terrestrial animals.[86] The blood has a high density of red blood cells, which contain oxygen-carrying haemoglobin. The oxygenated blood can be directed towards only the brain and other essential organs when oxygen levels deplete.[87][88][89] The spermaceti organ may also play a role by adjusting buoyancy (see below).[90] The arterial retia mirabilia are extraordinarily well-developed. The complex arterial retia mirabilia of the sperm whale are more extensive and larger than those of any other cetacean.[81]
If the killer whale pod is large, its members may sometimes be able to kill adult female sperm whales and can at least injure an entire pod of sperm whales.[200][201] Bulls have no predators, and are believed to be too large, powerful and aggressive to be threatened by killer whales.[202] Solitary bulls are known to interfere and come to the aid of vulnerable groups nearby.[203] However, the bull sperm whale, when accompanying pods of female sperm whales and their calves as such, may be reportedly unable to effectively dissuade killer whales from their attacks on the group, although the killer whales may end the attack sooner when a bull is present.[204][205] However, male sperm whales have been observed to attack and intimidate killer whale pods in competitive feeding instances. An incident was filmed from a long-line trawler: a killer whale pod was systematically taking fish caught on the trawler's long lines (as the lines were being pulled into the ship) when a male sperm whale appeared to repeatedly charge the killer whale pod in an attempt to drive them away; it was speculated by the film crew that the sperm whale was attempting to access the same fish. The killer whales employed a tail outward and tail-slapping defensive position against the bull sperm whale similar to that used by female sperm whales against attacking killer whales.[206] However, at some potential feeding sites, the killer whales may prevail over sperm whales even when outnumbered by the sperm whales. Some authors consider the killer whales \"usually\" behaviorally dominant over sperm whales but express that the two species are \"fairly evenly matched\", with the killer whales' greater aggression, more considerable biting force for their size and predatory prowess more than compensating for their smaller size.[204][207]
It is estimated that the historic worldwide population numbered 1,100,000 before commercial sperm whaling began in the early 18th century.[3] By 1880, it had declined by an estimated 29 percent.[3] From that date until 1946, the population appears to have partially recovered as whaling activity decreased, but after the Second World War, the population declined even further, to 33 per cent of the pre-whaling population.[citation needed] Between 184,000 and 236,000 sperm whales were killed by the various whaling nations in the 19th century,[240] while in the 20th century, at least 770,000 were taken, the majority between 1946 and 1980.[241]
The total number of sperm whales in the world is unknown, but is thought to be in the hundreds of thousands.[3] The conservation outlook is brighter than for many other whales. Commercial whaling has ceased,[3] and the species is protected almost worldwide, though records indicate that in the 11-year period starting from 2000, Japanese vessels have caught 51 sperm whales.[needs update] Fishermen do not target the creatures sperm whales eat,[3] but long-line fishing operations in the Gulf of Alaska have complained about sperm whales stealing fish from their lines.[164] 59ce067264